What is the Difference Between Protoxylem and Metaxylem?

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Protoxylem and metaxylem are two components of primary xylem, which is responsible for the transportation of water and minerals in plants. The main differences between protoxylem and metaxylem are:

  1. Formation: Protoxylem is the first formed primary xylem, maturing before the plant organs complete their elongation, while metaxylem is the later formed primary xylem, maturing after the completion of plant organ growth.
  2. Cell Size: Protoxylem has smaller cells, making it less prominent, whereas metaxylem has larger cells, making it more prominent.
  3. Lignification: Protoxylem has limited lignification, making it less effective in conducting water, while metaxylem has extensive lignification, making it more effective in conducting water.
  4. Water Conduction Efficiency: Metaxylem is more efficient in water conduction compared to protoxylem.
  5. Location: In a stem, protoxylem is located towards the outside, while metaxylem is located towards the inside.
  6. Cell Wall Thickenings: Protoxylem has annular or spiral thickenings around the vessels, while metaxylem has reticulate, scalariform, or pitted thickenings around the vessels.

In summary, protoxylem is the first primary xylem formed during plant growth and has smaller, less effective cells for water conduction, while metaxylem is the later formed primary xylem with larger, more effective cells for water conduction.

Comparative Table: Protoxylem vs Metaxylem

Here is a table summarizing the differences between protoxylem and metaxylem:

Feature Protoxylem Metaxylem
Formation Develops first during primary growth Develops after protoxylem during primary growth
Maturation Matures before plant organs complete their elongation Matures after the completion of growth of plant organs
Location In the stem, protoxylem is arranged towards the center In the stem, it is arranged towards the exterior
In the root, it is arranged towards the periphery In the root, it is arranged towards the interior
Cell Size Consists of smaller cells, leading to a narrow vessel lumen Consists of larger cells, leading to a wider vessel lumen
Lignification Less extensive lignification More extensive lignification
Efficiency Less efficient in water and mineral transport More efficient in water and mineral transport
Cell Types Contains more parenchyma cells Contains fewer parenchyma cells and more xylem fibers
Thickenings Annular or spiral thickenings are found in protoxylem The secondary cell walls of metaxylem vessels have scalariform, reticulate, and pitted thickenings
Fibers Fibers are absent in protoxylem Fibers are present in metaxylem
Tyloses Tyloses are not formed in protoxylem Tyloses may be present in metaxylem

Both protoxylem and metaxylem are primary xylem elements responsible for water and mineral conduction, and they contain living and dead cells. They develop from the primary meristem, pro-cambium, and contain tracheids, vessels, and parenchyma.